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The Form Is Never Final

Summary

This essay argues that the human form is not final: natural selection proceeds in a second-order, technogenic, natural environment that the species manufactures. Understanding "form" as an integrated configuration (body, cognitive and relational dispositions, technical-symbolic couplings), it is shown how the shift from the risk of early mortality to effective reproductive fitness reparameterizes evolution. In the technogenic niche, the informational mediation of the encounter (metric proximity, catalogue-algorithmic hierarchy-filter) favors assortativity and quantile matching; functional externalization redistributes cognitive costs; and mental health operates as a selective conduit, from extreme cutting to chronic friction that postpones parenthood. A verification program is proposed with comparable indicators (nulliparity, age at first birth, assortativity coefficients, prevalence of sexual inactivity), arguing that the data function as philosophical operators that make hair selection measurable. It is concluded that, by reconfiguring the environment, we reconfigure the pressures that shape us: the evolutionary future of humans is the multigenerational result of the ecologies we build.

Palavras‑chave: second grade natural environment; hair selection; effective reproductive fitness; metric proximity; quantile matching; functional externalization.

Initial delimitation

What we call the human form - bodily, psychic, relational and technical-symbolic - is not a static given nor a teleological end point. It is a local functional stability of the species, always negotiated with the environment. In this essay, the focus is on the evolutionary dynamics of the species under 2nd order selective forces - technically mediated forces that integrate the continuity of the natural. Here, "form" designates the integrated configuration of body, neurocognitive and affective processes, relational patterns, and technical-symbolic couplings that sustain reproduction and common life. Digressions on "incompleteness" in general or on ethical-political implications are avoided: the issue here is to describe, with material precision, how certain mechanisms of pairing, effective reproductive fitness and technical mediation reparametrize the evolution of the species. These dimensions are not compartments, but reciprocal couplings: variations in the body reorganise the psychic and relational circuit; changes in technical-symbolic mediation reparametrize cognitive dispositions and patterns of encounter.

It is important to distinguish two environmental regimes here: the first-degree natural environment - physicobiological - in which the species was configured throughout most of its evolutionary history, and the second-degree natural environment - the technogenic - which progressively thickens and today co-determines living conditions. It is not an "outside" of nature, but a layer manufactured by us, with its own rhythms, signs and accesses that rewrite opportunities for pairing and care, demanding new forms of attention, coordination and maintenance.

We call this technogenic environment a natural environment of the second degree - and not "artificial" - because it remains entirely within the material order: it is produced by natural agents (us), operates through the same physical, chemical and biological causalities and keeps intact the requirements of selection (variation, inheritance and differential reproduction). What changes is not the principle, but the configurations: new risk gradients, access and pairing. Therefore, there continues to be natural selection, now under manufactured conditions that co-evolve with us.

This reading finds support in contemporary lines of investigation: LeroiGourhan (1964; 1965) on technical externalization and gesture-language coupling; Simondon (1958; 1989) on technical individuation and "associated means"; the vehicular externalism of Clark and Chalmers (1998) and the Material Engagement Theory of Malafouris (2013), which treat cognition as a body-thing-signal system; and, at the evolutionary level, niche construction (OdlingSmee, Laland, and Feldman 2003; Laland 2016) and gene - culture coevolution (Richerson and Boyd 2005). They all converge on the idea that the technique integrates the continuity of the natural and reconfigures the conditions of selection.

The non-finality of the human form

By creating its own environment today, the species alters the pressures that select it. The development of a global technogenic niche reconfigures the selective landscape: the force of selection shifts mainly from early mortality to differences in effective reproductive fitness - that is, to variations in probability, timing and number of viable offspring. This shift explains why we place differential reproduction at the center: in evolution, differences that do not translate into viable offspring die with the individual. Only when variation, inheritance and differential reproduction converge does selection alter distributions (biological and symbolic-technical) over generations. It is, therefore, in the way you pair, when you pair and with whom you pair that today a large part of the evolutionary destiny of the species is decided.

This non-purpose has solid support both in the philosophy of biology and in contemporary biology. On a philosophical level, processualist proposals (Dupré and Nicholson 2018) reject the idea of ​​fixed forms and think of living beings as open processes, continually maintained and transformed by material and informational flows. At the empirical level, population genomics identifies extensive signals of recent and ongoing selection in humans (Akey 2009; Hawks et al. 2007), while demographic and longitudinal studies show ongoing selection in contemporary populations: direct estimates in cohorts like Framingham (Byars et al. 2010), evidence of microevolutionary change in ages at first birth in a historical context (Milot et al. 2011), and relationships between patterns reproductive and polygenic profiles (Beauchamp 2016; Kong et al. 2017). Taken together, these lines maintain that human evolution continues, without teleology, and that "form" - understood here as an integrated configuration - remains open to selective reconfigurations imposed by an increasingly technogenic environment.

'Form' in this essay is more than morphology: it includes cognitive and relational dispositions that emerge from material couplings with the environment and with artefacts.

2nd order forces and the technogenic niche

Before mapping these forces, it is important to note a historical point: the species did not always manufacture its environment in a systematic way. For most of its evolutionary history, the relevant environment was mainly the first degree. The regular creation of the environment by the species itself becomes more dense with agriculture and sedentarization (control of cycles, storage, division of labor) and then accelerates with industry and modern technical networks. It is this process that prepares the second-order natural environment in which contemporary selective pressures operate. In this key, the demographic transition (Omran 1971) and niche construction readings help to clarify that selection did not stop: it was reconfigured as the medium also began to be manufactured.

Technique is not external to life: it is a secondary level of the natural, a material continuity that co-determines the environment. In this light, we can distinguish four families of forces that together reparametrize costs and benefits of survival and reproduction.

Biotechnical strengths include vaccination, advanced obstetrics and neonatology, assisted reproduction, and gene screening/editing. Its clearest effect was to drastically reduce pre-reproductive mortality, shifting selection towards traits that favor pairing and parental care and altering ages and reproductive trajectories. It is the classic terrain of evolutionary medicine (Nesse 2010; Stearns 1999) and recent population genomics, where ongoing selection is estimated even in contexts with strong medical intervention; the demographic transition (Omran 1971) precisely describes this shift in risks that reparameterizes fitness.

Technoecological forces derive from urbanization, chronic pollutants, controlled microenvironments and climate change of anthropogenic origin. They produce low-intensity, persistent pressures that require distinct immunoendocrine and metabolic coordination and decouple physical effort from survival, favoring profiles capable of persisting in materially complex environments. Concepts such as exposome (Wild 2005) and urban metabolism (Wolman 1965; Bettencourt 2013) help name this web of exposures and artificial flows that make up the second degree of the medium; the planetary health agenda (Whitmee et al. 2015) shows how these manufactured conditions return as biological constraints.

Infosymbolic forces impact directly on the encounter: matching platforms and catalog systemsalgorithmic hierarchyfilter convert physical proximity into metric proximity. The apparent abundance of options, the idealization of images and the segmentation of niches raise acceptance thresholds, generating choice overload and an expanded assortativity. The typical result is a densification of intraquantile pairings (by education/status, by signs of attractiveness, by height), with effects on the timing and stability of reproductive links. Here, cultural evolution (Henrich 2016) provides the framework for understanding learning biases and preferences; studies on line matching and homogamy (Rosenfeld, Thomas, and Hausen 2019; Kalmijn 1998; Schwartz 2013) show how filters magnify homophily; and the literature on choice overload (Iyengar and Lepper 2000) helps to read postponement and indecision as products of saturated catalogs and permanent algorithmic hierarchies.

In the infosymbolic family, language and image function as signaling and screening operators: discursive and image patterns stabilized by recommendation algorithms modulate attention, expectation and acceptance. By converting preferences into comparable signals, the semiotic ecology of the encounter reconfigures search costs, raises acceptance thresholds and thickens assortativity, without breaking the material continuity of the process.

Cyborg forces encompass prosthetics, implants, psychotropic drugs, brain-machine interfaces (BCI) and exoskeletons. They create performance differentials that translate into indirect effects on income, social network and longevity, influencing the reproductive contribution throughout life. The philosophy of mind and technique has a precise formulation here: NaturalBorn Cyborgs (Clark 2003) and Simondon's technical individuation describe these couplings as stable operative extensions, not as props; Research in BCI and neuroprosthetics shows how such extensions can crystallize new functional margins.

How the manufactured medium translates into selection

With the drop in infant and juvenile mortality, selection acts mainly in adulthood (Omran 1971). In operational terms, this shifts the relevant variation to traits that affect who can form reproductive pairs, when they do so, and with what stability - exactly what evolutionary demography has sought to measure through restrictions on fertility and survival calendars.

In other words: selection now increasingly focuses on the ability to operate in the natural environment of second degree. On average, profiles with greater resistance to stress, tolerance to social pressure and competence for sustained cognitive effort tend to persist - that is, those who manage to maintain profiles capable of sustaining divided attention, working memory and decision making under permanent comparison, without operational breakdown. In parallel, robust affective regulation is required in the face of evaluation anxieties and social comparison scales which, when persistent, degrade the margin of self-modulation and reduce effective reproductive fitness. What changes, in depth, is the structure of the pairing: from the local meeting we move to the meeting through measured affinities. Online dating displaces traditional intermediaries and becomes the dominant way of meeting (Rosenfeld, Thomas and Hausen 2019), at the same time as it reveals hierarchies of desirability and strong response gradients, with a sharp drop in the probability of message return as the distance in the algorithmic hierarchy between pursued and pursuer increases (Bruch and Newman 2018). In saturated catalogs, choice overload and permanent algorithmic comparison raise acceptance thresholds and encourage postponement (Iyengar and Lepper 2000).

This results in a tendency towards quantile matching, which increases the covariance between traits in couples and stratifies the distribution of resources and cultural capital in descendants. In terms of quantitative genetics, phenotypic assortativity increases additive variance and reinforces intergenerational similarities (Lynch and Walsh 1998). However, recombination and regression to the mean function as statistical brakes in the short and medium term, maintaining porosity between subpopulations (Lynch and Walsh 1998). Hence the image of a reproductive archipelago: panmixia is diluted in global subnetworks with soft barriers to gene flow, without implying speciation.

The functional externalization of memory, orientation, and coordination out of the biosome redirects costs from what the body needs to maintain internally and shifts capabilities toward coordinated artifacts, routines, and environments - it is the lesson of distributed cognition (Hutchins 1995) and cognitive artifacts (Norman 1991), compatible with vehicular externalism (Clark and Chalmers 1998). These technical-symbolic couplings are not props: they condition performance, support networks and, therefore, reproductive trajectories.

Mental health functions here as a decisive channel on two levels: at the extreme end, suicide implies zero aptitude; in chronic friction, persistent depression reduces the likelihood of conception and delays first birth, with recent findings in couple samples and population registries (Liao 2024; Golovina 2023). These are capillary, cumulative effects, without moral teleology, but with a measurable evolutionary impact if they persist over time.

Empirical anchors and verification horizon

The recent empirical picture reinforces the view that the manufactured environment reconfigures effective reproductive fitness through capillary pathways. In highly digitized populations, increases in sexual inactivity and reproductive delay have been consistently observed. In the US, sexual inactivity increased between 2000 and 2018, especially in men 18 - 24 and 25 - 34, with about one in three men 18 - 24 not having sexual activity in the last year (Ueda et al. 2020), converging with the drop in sexual frequency documented in GSS series (Twenge, Sherman, and Wells 2017). In Japan, successive analyzes of the National Fertility Survey show high and increasing levels of sexual inexperience among 18 - 39 year olds and, among those 18 - 34 never married, more than 40% without heterosexual experience in 2015 (Ghaznavi et al. 2019); the 2021 round confirms objective obstacles to the encounter and decline of marriage (NIPSSR 2022). These patterns are compatible with the displacement of intermediaries and the dominance of online encounters (Rosenfeld, Thomas, and Hausen 2019), suggesting that metric proximity and filters reinforce postponement and assortativity.

Mental health emerges as a measurable selective channel. In cohorts of couples from the general population, preconception depression is associated with longer time to conception and increased risk of infertility (Liao et al. 2024). In national registries, depression treated in secondary care is associated with a lower probability of having children and lower total fertility, with differences by sex and socioeconomic status (Golovina et al. 2023); Close results appear when modeling own depression, peer depression, and childlessness (KailaheimoLönnqvist et al. 2024). These findings do not establish fatalism; They only indicate chronic friction with reproductive effects when they persist.

In this way, the thesis is verifiable by a program of indicators and comparable series: - Nulliparity by cohort and by subgroups of exposure to the technogenic niche (education, income, urbanization), using standardized sources (Human Fertility Database 2025; see also Jasilioniene et al. 2016). - Age at first birth by status/education quantiles and distribution of interval to first birth, to detect timing frictions (Human Fertility Database 2025). - Assortativity coefficients in couples (education/status), tracking the tendency of reinforced homophily in digitalized ecologies (Kalmijn 1998; Schwartz 2013) and its covariance with the mediatization of the encounter (Rosenfeld, Thomas and Hausen 2019). - Prevalence of sexual inactivity/inexperience by age and sex in national surveys (Ueda et al. 2020; Ghaznavi et al. 2019) and its relationship with digital mediation indicators. - Mental health and parenting: relative risk of nulliparity and time to conception by own and partner depression profiles (Liao et al. 2024; Golovina et al. 2023; KailaheimoLönnqvist et al. 2024).

Environmental contingency is part of the philosophical and empirical proof: if policies that reduce the material costs of parenting and infrastructures that facilitate non-metric encounters reduce involuntary celibacy or bring forward the first child, then the selective process operates through the environment we make - not through immutable essences. It is this differential response, measured in decades and compared in cohorts, that confirms 2nd order selection.

These indicators are philosophical operators, not just illustrations: they function as materialist proxies of reproductive costs and benefits under the technogenic niche. Their reading requires causal caution (context, cohorts, heterogeneity), but it is precisely this contingency that makes them probative for a 2nd degree selection.

Conclusion

What has been established is simple and operative: the human form is not final because natural selection continues to act, today especially in the second-grade, technogenic natural environment, which the species itself manufactures. The form, understood as an integrated configuration of the human, is therefore a local stability subject to reparameterizations whenever the environment changes.

The originality of this proposal does not lie in claiming an isolated discovery, but in articulating four threads that rarely appear together: a materialist ontology without dualisms between nature and technique; the distinction between first and second degree means; the centrality of differential reproduction as a criterion for effective change; and the translation of the effects of digital mediation into nameable mechanisms - metric proximity, funnel effect, quantile matching, reproductive archipelago - with verifiable metrics.

A practical consequence follows from this framework: there is no fixed destination. By reorganising the environment, we reorganise the selective pressures that shape us. Policies, infrastructures and habits can attenuate or intensify trajectories - and this is precisely what series and cohorts will allow us to measure.

Finally, the limit is recognized: these are slow trends, dependent on the stability of the technogenic niche and heterogeneous by context. The opening of the work is to follow the evidence and test the predictions; philosophical openness, in maintaining vigilant thinking about the compatibilities that our own environment imposes.

Thinking about policies and infrastructures is, here, describing ways of modulating the niche: material changes to the environment that redistribute reproductive costs and benefits. It's not moralizing; is to make explicit the selective pragmatics of the ecologies we manufacture.

References

Akey, Joshua M. 2009. "Constructing Genomic Maps of Positive Selection in Humans: Where Do We Go from Here?" Genome Research 19 (5): 711 - 22.

Beauchamp, Jonathan P. 2016. "Genetic Evidence for Natural Selection in Humans in the Contemporary United States." Proceedings of the National Academy of Sciences 113 (28): 7774 - 79.

Bettencourt, Luís M. A. 2013. "The Origins of Scaling in Cities." Science 340 (6139): 1438 - 42.

Bruch, Elizabeth E., e M. E. J. Newman. 2018. "Aspirational Pursuit of Mates in Online Dating Markets." Science Advances 4 (8): eaap9815.

Clark, Andy. 2003. Natural-Born Cyborgs: Minds, Technologies, and the Future of Human Intelligence. Oxford: Oxford University Press.

Clark, Andy, e David J. Chalmers. 1998. "The Extended Mind." Analysis 58 (1): 7 - 19.

Dupré, John, e Daniel J. Nicholson, eds. 2018. Everything Flows: Towards a Processual Philosophy of Biology. Oxford: Oxford University Press.

Ghaznavi, Chandni, Haruka Sakamoto, Daisuke Yoneoka, Shoko Nomura, Kenji Shibuya, e Peter Ueda. 2019. "Trends in Heterosexual Inexperience among Young Adults in Japan: Analysis of National Surveys, 1987 - 2015." BMC Public Health 19: 355.

Golovina, Anastasia, e col. 2023. "Depression and Completed Fertility: Evidence from National Registers." Demography 60 (4): 1201 - 23.

Hawks, John, Eric T. Wang, Gregory Cochran, Henry C. Harpending, e Robert K. Moyzis. 2007. "Recent Acceleration of Human Adaptive Evolution." Proceedings of the National Academy of Sciences 104 (52): 20753 - 58.

Henrich, Joseph. 2016. The Secret of Our Success: How Culture Is Driving Human Evolution, Domesticating Our Species, and Making Us Smart. Princeton, NJ: Princeton University Press.

Human Fertility Database (HFD). 2025. Human Fertility Database. Max Planck Institute for Demographic Research e Vienna Institute of Demography. https://www.humanfertility.org.

Hutchins, Edwin. 1995. Cognition in the Wild. Cambridge, MA: MIT Press.

Iyengar, Sheena S., e Mark R. Lepper. 2000. "When Choice Is Demotivating: Can One Desire Too Much of a Good Thing?" Journal of Personality and Social Psychology 79 (6): 995 - 1006.

Jasilioniene, Aiva, Dmitri A. Jdanov, Tomáš Sobotka, e col. 2016. "Data Resource Profile: The Human Fertility Database." International Journal of Epidemiology 45(4): 1077 - 85.

KailaheimoLönnqvist, Sini, e col. 2024. "Depression, Partner's Depression, and Childlessness: A Population Register Study." Social Psychiatry and Psychiatric Epidemiology 59 (2): 345 - 58.

Kalmijn, Matthijs. 1998. "Intermarriage and Homogamy: Causes, Patterns, Trends." Annual Review of Sociology 24: 395 - 421.

Kong, Augustine, e col. 2017. "Selection against Variants in the Genome Associated with Educational Attainment." Proceedings of the National Academy of Sciences 114 (5): E727 - E732.

Laland, Kevin N. 2017. Darwin's Unfinished Symphony: How Culture Made the Human Mind. Princeton, NJ: Princeton University Press.

Leroi-Gourhan, André. 1964. The Gesture and the Word. Flight. 1, Technique and Language. Paris: Albin Michel.

- - - . 1965. The Gesture and the Word. Flight. 2, Memory and Rhythms. Paris: Albin Michel.

Liao, Tong, e col. 2024. "Preconception Depression Reduces Fertility: A Couple-Based Study." Human Reproduction Open 2024 (3): hoae032.

Lynch, Michael, e Bruce Walsh. 1998. Genetics and Analysis of Quantitative Traits. Sunderland, MA: Sinauer.

Malafouris, Lambros. 2013. How Things Shape the Mind: A Theory of Material Engagement. Cambridge, MA: MIT Press.

National Institute of Population and Social Security Research (NIPSSR). 2022. The 14th Japanese National Fertility Survey: Marriage Process and Fertility of Married Couples, Attitudes toward Marriage and Family among Japanese Singles. Tokyo: NIPSSR.

Norman, Donald A. 1991. "Cognitive Artifacts." In Designing Interaction: Psychology at the Human-Computer Interface, editado por John M. Carroll, 17 - 38. Cambridge: Cambridge University Press.

Odling-Smee, F. John, Kevin N. Laland, e Marcus W. Feldman. 2003. Niche Construction: The Neglected Process in Evolution. Princeton, NJ: Princeton University Press.

Omran, Abdel R. 1971. "The Epidemiologic Transition: A Theory of the Epidemiology of Population Change." The Milbank Memorial Fund Quarterly 49 (4): 509 - 38.

Richerson, Peter J., e Robert Boyd. 2005. Not by Genes Alone: How Culture Transformed Human Evolution. Chicago: University of Chicago Press.

Rosenfeld, Michael J., Reuben J. Thomas, e Sonia Hausen. 2019. "Disintermediating Your Friends: How Online Dating in the United States Displaces Other Ways of Meeting." Proceedings of the National Academy of Sciences 116 (36): 17753 - 58.

Schwartz, Christine R. 2013. "Trends and Variation in Assortative Mating: Causes and Consequences." Annual Review of Sociology 39: 451 - 70.

Simondon, Gilbert. 1958 (1989). Du Mode d'Existence des Objets Techniques. Paris: Aubier.

Twenge, Jean M., Ryne A. Sherman, e Brooke E. Wells. 2017. "Declines in Sexual Frequency among American Adults, 1989 - 2014." Archives of Sexual Behaviour 46 (8): 2389 - 2401.

Ueda, Peter, Naomi M. Means-Christensen, Ryne A. Sherman, e Jean M. Twenge. 2020. "Trends in Frequency of Sexual Activity and Number of Sexual Partners among Adults in the United States, 2000 - 2018." JAMA Network Open 3 (6): e203833.

Whitmee, Sarah, e col. 2015. "Safeguarding Human Health in the Anthropocene Epoch: Report of The Rockefeller Foundation - Lancet Commission on Planetary Health." The Lancet 386 (10007): 1973 - 2028.

Wild, Christopher P. 2005. "Complementing the Genome with an 'Exposome': The Outstanding Challenge of Environmental Exposure Measurement in Epidemiology." Cancer Epidemiology, Biomarkers & Prevention 14 (8): 1847 - 50.

Wolman, Abel. 1965. "The Metabolism of Cities." Scientific American 213 (3): 179 - 90.